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Table 2 Proportions of variance components influencing the individual birth weight of puppies. Comparison of the variance components of a model without covariates (1) and a model with covariates (2). The direct-maternal covariance was held at zero

From: Heritabilities for the puppy weight at birth in Labrador retrievers

Covariate/Variance

Model 1

Model 2

Heritability direct

0.21 ± 0.047

0.17 ± 0.041

Heritability maternal

0.22 ± 0.053

0.22 ± 0.049

Dam environmental proportion

0.07 ± 0.034

0.07 ± 0.031

Litter environmental proportion

0.14 ± 0.013

0.08 ± 0.010

Residual proportion

0.36 ± 0.033

0.46 ± 0.032

Total heritability*

0.32

0.28

  1. * \( {h}_T^2=\left({\sigma}_d^2+0.5\bullet {\sigma}_m^2+1.5\bullet {\sigma}_{dm}\right)/{\sigma}_p^2 \)